I would say from what you have told me that the problem was too little humidity when drying. Gotta keep it on the 40% so that the weed don't dry too fast and in a bad way. But the buds are looking really good so you sure they are bad?
Good luck with your grow I hope you get the best taste of buds and healthy plants! Take care and stay lit! I will drop by sometimes to check on your process! Say hello if you ever visit my page!
Greetings from L.S.T ππΌπ