I think your light is too strong. Even if it took a while to see symptoms.
Without supplemental co2, you're better off with ~65-70% power, 18" and 18 hour cycle.
if your heart is set on 20 hours, reduce proportionally. ~55% for 20 hour operation.
Also possible that your 2tbsp/week is not fully supplementing the soil, but this looks to be localized to the tallest parts of plants and not the short one much at all, plus personal familiarity with the fc4800. Being this little off might take weeks to see a symptom caused by it.