Yes, both on, even during the dark cycle.
But you can get away with running your fans for only15 minutes every hour while in early to mid veg, but having them on 100% of the time during flowering is mandatory.
I usually have em on a timed switch (smart plug), and have them running for 15 minutes every hour during the day. At night, I have them on 15 minutes every 2hrs. Make sure not pointing them at the plants directly.