Leaf releasing moisture faster than it uptakes. Plant chooses which parts of leaf to "sacrifice" by choosing the least vital parts of a leaf to be cannabalized first.
This is caused by a breakdown in water system and the cycle as a whole. Where the bottleneck is, could be a number of things. The plant actively manages balance primarily through stomatal control, to ensure that the rate of water uptake from the soil can keep pace with the rate of water loss to the atmosphere, thus preventing lethal levels of water stress. The entire system is a balance between the atmospheric "demand" for water and the physical limitations and resistances within the soil and the plant itself.
Soil/Roots,
Plant,
Air/Atmosphere,
I could waffle on forever, find where the bottleneck is and why water stopped moving.