They are ripeening.
genetic variety. not all plants can grow big buds. Yes, your environment and choices can impact consistency of meeting potential, but not nearly as much as most perceive.
Does look a bit overfed, could be N, but more than one thing causes dark foliage. The glossiness helps point the finger at N, too, but could be more than one thing building up. Being a little overfed i not the cause of your bud sizes, though. It's not catastrophically overfed or anything. There may be a measurable correlation to a negative effect but it is not gauranteed to be profound. Some plants will be more robust to mistreatment than others and still nearly reach potential. I've had plenty of slightly dark foliage still result in a proper yield. When that darkness creeps in, back off sooner next time.
g/sq ft is a better metric to look at than bud size. If getting 50g/sq ft or more in ambient co2 conditions, you probably didn't 'lose' too much. As longn as canopy is full, yield will come. How yo managed that canopy will impact bud size too. More branches = smaller, more distributed fower mass. you only need about 3 colas per sq ft to maximize yield.