some deficiency that FF doesn't cover... looks like Ca or Mg, i'm forgetting the difference at moment. think that's Ca
pH could be a culprit, too - it can cause any and all def and tox. So, if you have been bombing it with feed or if pH is off, back off for a while and see if the problem abates on its own. then, you can think about doing something more active with a bit more confidence.